H1' C1' N9 C4 : chi adenine H1' C1' N9 C8 : chi adenine C4' C3' O3' +P : epsilon C2' C3' O3' +P : epsilon H3' C3' O3' +P : epsilin P O5' C5' C4': beta H4' C4' C5' H5': Gamma, averaged only on Yokoyama1 and Yokoyama2 H3' C3' C4' H4': Delta, averaged only on Yokoyama1 and Yokoyama2